Dr Kim Last

        Head and shoulder photo of a windswept Dr Kim Last on a Scottish beach

Marine Biologist

i am a biologists interested in the behaviour of marine organisms specifically their response to environmental change.

Contact details:
  • kim.last@sams.cxyb888.cn
  • +44 (0)1631 559 414

  • clockworkworm

i am interested in the effects of human activities on marine organisms at behavioural and physiological levels.

七星彩走势图带坐标准版as humans change the face of our planet so animals have to adapt or go extinct – i hope even in a small way to help avoid the latter.

七星彩走势图带坐标准版underlying my work is a fascination of clock biology or chronobiology i.e. how animals tell the time. this has led me from studying the chronobiology of worms in muddy estuaries to zooplankton across the entire arctic ocean – i have surprisingly found that both demonstrate an uncanny knack for being in the right place at the right time.  

七星彩走势图带坐标准版i see myself as a 'practical' scientist and enjoy answering difficult questions by developing new technologies, often in very hostile places, in order to better link the biology with the physics of the natural environment.



Arctic PRIZE: Climate change impacts in the Arctic Ocean. Funded by NERC. 2016-19

DIAPOD七星彩走势图带坐标准版: Climate change impacts on marine copepods. Funded by NERC. 2016-19

Arctic ABC七星彩走势图带坐标准版: Arctic marine ecology and autonomous technology development. Funded by NFR. 2014-18

CACHE七星彩走势图带坐标准版: Climate change impacts on commercial shellfish. Funded by EU FP7. 2013-17

Nephrops Survival 2



: circadian rhythms in arctic ecosystems. funded by nfr. 2012-15

七星彩走势图带坐标准版tromyt projects (3): effects of seawater chlorination on mussels. funded by cefas/edf. 2013-17

SabALV: Effects of seawater chlorination on Sabellaria polychaetes. Funded by CEFAS/EDF. 2011

maree: sediment impacts from marine renewables on commercial and biogenic reef builders. funded by hie/eu/sfc. 2010-14

七星彩走势图带坐标准版malf: impacts of aggregate dredging on benthic organisms. funded by defra. 2007-10


Hutchison ZL, Green DH, Burrows MT, Jackson AC, Wilson B, Last KS (2020) Survival strategies and molecular responses of two marine mussels to gradual burial by sediment. Journal of Experimental Marine Biology and Ecology 527: 151364  

Stewart-Sinclair PJ, Last KS, Payne BL, Wilding TA (2020) A global assessment of the vulnerability of shellfish aquaculture to climate change and ocean acidification. Ecology and Evolution七星彩走势图带坐标准版 10(7): 3518–3534.  

Geoffroy M, Berge J, Majaneva S, Johnsen G, Langbehn TJ, Cottier F, Mogstad AA, Zolich A, Last K (2018) Increased occurrence of the jellyfish Periphylla periphylla in the European Arctic. Polar Biology 41(12): 2615-2619

Daase M., Kosobokova   K., Last K. S., Cohen J., Hatlebakk M., Søreide J., Choquet M. 2018.  New insights into the biology of Calanus spp. (Copepoda) males in the Arctic. Marine Ecology Progress Series. DOI: .

Häfker, N. S., Teschke, M., Last, K. S., Pond, D. W., Hüppe, L., Meyer, B., 2018. Calanus finmarchicusseasonal cycle and diapause in relation to gene expression, physiology, and endogenous clocks. Limnology and Oceanography 9999: 1-24. DOI: https://doi.org/10.1002/lno.11011

Benjamins, S. Fox, C. J., Last, K. S., McCarty, C. E., 2018. Individual identification of flapper skate Dipturus intermedius using a baited camera lander. Endangered Species Research 37, p37.
Hobbs, L., Cottier, F., Last, K. S., Berge, J. 2018. Pan-Arctic diel vertical migration during the polar night. Marine Ecology Progress Series.

Sören Häfker, Bettina Meyer, Kim S. Last, David W. Pond, Lukas Hüppe, Mathias Teschke, 2017. Circadian clock involvement in zooplankton diel vertical migration. Current Biology. DOI:

Kent, F.E., Gray, M.J., Last, K.S., Sanderson, W.G., 2016. Horse mussel reef ecosystem services: evidence for a whelk nursery habitat supporting a shellfishery. International Journal of Biodiversity Science, Ecosystem Services & Management, 1-9.

Orejas, C., Gori, A., Rad-Menéndez, C., Last, K. S., Davies, A. J., Beveridge, C. M., Sadd, D., Kiriakoulakis, K., Witte, U., Roberts, J. M. 2016. The effect of flow speed and food size on the capture efficiency and feeding behaviour of the cold-water coral Lophelia pertusa. JEMBE 481, 34-40.

Last, K. S., Hendrick, V. J., Beveridge, C. M., V. J., Wilding, T., Roberts, D. A. Lethal and sub-lethal responses of the biogenic reef forming polychaete Sabellaria alveolata to aqueous chlorine and temperature. Marine Environmental Research. 117: 44–53.

Hutchison, Z. L., Hendrick, V. J., Burrows, M. T., Wilson, B., Last, K. S. Buried Alive: The behavioural response of two mussels, Modiolus modiolus and Mytilus edulis to sudden burial by sediment. PLoS ONE. http://dx.doi.org/10.1371/journal.pone.0151471

Hendrick, V.J., Hutchison, Z., Last, K. S. Sediment burial intolerance of marine macroinvertebrates. PLoS ONE. 11, e0149114.

Cottrell, R. S., Hutchison, Z. L., Black, K., Last, K. S. 2016. The influence of organic material and temperature on the burial tolerance of the blue mussel, Mytilus edulis: considerations for the management of marine aggregate dredging. PLoS ONE. DOI:10.1371/journal.pone.0147534

Last, K. , Hobbs, L., Berge, J., Brierley, A. S., Cottier, F. 2016. Moonlight drives ocean-basin scale mass vertical migration of zooplankton during the Arctic winter. Current Biology. 26: 1-8.

Berge, J., Daase, M., Renaud, P., Ambrose, W., Darnis, G., Last, K.S.,七星彩走势图带坐标准版 Leu, E., Cohen, J., Johnsen, G., Moline, M.A., Cottier, F., Varpe, Ø., Shunatova, N., Balazy, P., Morata, N., Massabuau, J.-C., Falk-Petersen, S., Kosobokova, K., Clara J.M. Hoppe, Weslawski, J.M., Kuklinski, P., Legeżyńska, J., Nikishina, D., Cusa, M., Kedra, M., Wlodarska, M., Vogedes, D., Camus, L., Tran, D., Michaud, E., Gabrielsen, T.M., Granovitch, A., Gonchar, A., Krapp, R., Callesen, T.A., 2015 Unexpected levels of biological activity during the polar night offers new perspectives on a warming Arctic. Current Biology 25, 1-7.

Berge, J., Renaud, P.E., Darnis, G., Cottier, F., Last, K.S., Gabrielsen, T.M., Johnsen, G., Seuthe, L., Weslawski, J.M., Leu, E., 2015 In the dark: a review of ecosystem processes during the Arctic polar night. Progress in http://dx.doi.org/10.1016/j.pocean.2015.08.005

Cohen, J.H., Berge, J., Moline, M.A., Sørensen, A.J., Last, K.S七星彩走势图带坐标准版, Falk-Petersen, S., Renaud, P.E., Leu, E.S., Grenvald, J., Cottier, F., 2015 Is ambient light during the high Arctic Polar Night sufficient to act as a visual cue for zooplankton? PLoS ONE 10 (6), e0126247.

Browne, N. K., Precht, E., Last, K. S., Todd, P. A. 2014. The photo-physiological costs associated with acute sediment stress events in three near-shore turbid water corals. MEPS 502: 129-14.

Lai, S., Gillis, L. G., Mueller, C.,  Bouma, T. J., Guest, J. R., Last, K. S., Ziegler, A. D., Todd, P. A. 2013 First experimental evidence of corals feeding on seagrass matter. Coral Reefs. 32: 1061-1064.

Miller, R. G., Hutchison Z. L., Macleod, A. K., Burrows, M. T., Cook, E. J., Last K. S七星彩走势图带坐标准版., Wilson, B. 2013 Marine renewable energy development: assessing the Benthic Footprint at multiple scales. Frontiers in Ecology and the Environment. 11: 433-440.

Schaum CE, Batty R, Last KS (2013) Smelling danger – alarm cue responses in the polychaete Nereis (Hediste) diversicolor (Müller, 1776) to potential fish predation. PLoS ONE 8(10): e77431. doi:10.1371/journal.pone.0077431



Last, K. S., A clockwork world. Ocean Explorer Magazine, issue 37, summer 2017.

Last, K. S., 七星彩走势图带坐标准版 Book review of “Moonstruck: how lunar cycles affect life” by Ernest Naylor. Ocean Challenge, Vol 22, No 1.

Last, K. S., Beveridge, C. M. & Wilding, T. A. 2016. TROMyt4 – Tolerance of Mytilus edulis七星彩走势图带坐标准版 to seawater chlorination and suspended sediment. A report by SRSL for Cefas 02044_000, pp. 41.

Last, K. S., Beveridge, C. M. & Wilding, T. A. 2016. TROMyt3 A – Chlorine demand trial. A report by SRSL for Cefas, pp.19

Last, K. S., Beveridge, C. M. & Wilding, T. A. 2016. TROMyt3 B - Tolerance of Mytilus edulis to suspended sediment. A report by SRSL for Cefas, pp.25

Last, K. S., Beveridge, C. M. & Wilding, T. A. 2016. TROMyt3 C – Bromoform trial. A report by SRSL for Cefas, pp.22

Last, K. S., Hendrick, V. J. 2014. Snowy and Penny’s Arctic Adventure. Published by Sole Design.

Last, K. S., 七星彩走势图带坐标准版Hendrick, V. J. 2014 In “Annual, Lunar and Tidal Clocks”, chapter “The clock-work-worms: diversity and function of clock expression in marine polychaete worms.” Springer.

Last, K. S., Hendrick, V. J. 2014 “Predicted sensitivity of benthic organisms within the Don Diego Phosphate Marine Dredge Concession Area”. Report prepared on behalf of SRSL for Odyssey Marine Exploration.

Last, K. S., Beveridge, C. M., Wilding, T. 2014. “Pilot study on the tolerance of Mytilus edulis to aqueous chlorine and suspended sediments.” Commercial report for CEFAS BEEMS 2013, WP8. SRSL report No 01018_P06870001. 25pp

Last, K. S., Hendrick, V. J, Beveridge, C. M. 2012 Tolerance of Sabellaria alveolata to Aqueous Chlorine. Final Report for the BEEMS programme, Work Package 6.


七星彩走势图带坐标准版i have led a number of commercial projects over the last ten years with a value totaling >£600k. initial work focused on assessing the consequences of anthropogenic sediments from aggregate dredging on benthic macroinvertebrates (funded by the malsf programme). as part of this a novel mesocosm was built, the vortex resuspension tank (vorts), with the capacity to simulate sediment plumes. using this system two further projects (sabspin / sabalv) were commissioned, this time assessing the impacts of seawater chlorination on sensitive and protected biogenic reef habitats. following this, four further projects (tromyt1-4) were commissioned by cefas/edf energy all aimed at determining the impact of chlorination/sediment on the fouling blue mussel. collectively these projects have provided highly relevant and scientifically robust data to the marine aggregate industry (where the uk is the largest producer in the world) and a multi-billion pound new uk nuclear power station development. the vort mesocosms were also used as part of a large multi-partner study (maree) to assess the impacts of sediment from the marine renewable industry.



七星彩走势图带坐标准版sams consultancy work is delivered through its wholly owned commercial subsidiary, . (srsl). srsl operates a comprehensive quality management system (qms) that is accredited to iso9001:2008 standard, guaranteeing that all projects deliver the best possible service to our customers. all srsl projects are planned by a professional programme manager who oversees a team of dedicated project managers, all of whom are focussed specifically on client delivery, and have scientific experience and technical expertise in fields relevant to their assigned projects.

for further information concerning commercial enterprise opportunities at sams, please contact srsl by email () or phone +44 (0) 1631 559 470.


Marine Science BSc: Module leader for year 4 'Behaviour and Biological Clocks'

marine science bsc: lecturer year 4 'deep-sea ecosystems'

七星彩走势图带坐标准版marine science bsc: lecturer year 4 'polar ecology'

marine science bsc: lecturer year 3 'marine conservation'

七星彩走势图带坐标准版marine science bsc: lecturer year 2 'marine biology'


七星彩走势图带坐标准版in the last few years i have also contributed to various external undergraduate lectures and seminars, highlights include: environmental genomics masterclass at the university of newcastle; chronobiology at the university of svalbard (unis); sediment impact studies at the universities of singapore, stirling and bordeaux.

七星彩走势图带坐标准版i have also examined 5 phd and 1 msc theses.



Kati Michalek七星彩走势图带坐标准版: Climate change and shellfish. PhD 2014-17. University of Aberdeen



七星彩走势图带坐标准版laura hobbs: arctic ocean zooplankton migrations. phd 2012-16. university of aberdeen

flora kent: ecosystem services of horse mussel beds. phd 2012-15. heriot-watt university

zoe hutchison: sediment impacts on mussels. phd 2010-14. university of aberdeen

beatiz de francisco: ocean acidification in cold-water corals. phd 2007-13. university of aberdeen


tammi warrender: microbial associations of cold-water corals. msc 2014-15. university of st andrews

七星彩走势图带坐标准版ffion lowe: ocean acidification in cold-water corals. msc 2013-14. university of st andrews

七星彩走势图带坐标准版richard cottrell: sediment and organic matter impacts on mussels. msc 2013-14. university of st andrews

keshtav ramdial: metal leachate toxicity in marine systems. msc 2013-14. university of st andrews

natasha hill: ocean acidification in cold-water corals. msc 2013-14. university of st andrews

七星彩走势图带坐标准版fraser mackay: sediment impacts on feeding in cold-water corals. msc 2012-13. university of st andrews

七星彩走势图带坐标准版harriet condie: sediment impacts on sabellaria. msc 2008-09. york university

charlotte elisa schaum: fish alarm signals. msc 2008-09. university of rostock


moonlight helps plankton escape predators during arctic winters ()

do blame it on the moonlight: arctic plankton boogie to the moon ()

七星彩走势图带坐标准版tiny marine critters migrate in the arctic winter using moonlight ()

arctic winter: marine creatures migrate by moonlight during arctic winter, study says ()

七星彩走势图带坐标准版moonlight drives zooplankton’s winter migrations ()

七星彩走势图带坐标准版arctic animals navigate by light of the winter moon ()

七星彩走势图带坐标准版'werewolf' creatures in the arctic navigate by the cycle of the moon: during dark winter months plankton migrate in lunar cycles ()

artico, animali marini stregati da luna ()

七星彩走势图带坐标准版in arctic winter, marine creatures migrate by the light of the moon ()

marine creatures migrate by moon's light during arctic winter ()

biologists make surprise discovery among plankton in the arctic ()

'werewolf' creatures in the arctic navigate by the cycle of the moon: during dark winter months plankton migrate in lunar cycles ()

'werewolf' creatures in the arctic navigate by the cycle of the moon: during dark winter months plankton migrate in lunar cycles ()

moonlight drives marine creatures in arctic winter ()

arctic animals navigate by the light of winter moon ()

arctic animals navigate by the light of winter moon: study ()

七星彩走势图带坐标准版arctic animals navigate by the light of winter moon ()

moonlight drives marine creatures in arctic winter ()

七星彩走势图带坐标准版moonlight drives mass migration in arctic winter ()

七星彩走势图带坐标准版during arctic winters, active marine creatures migrate by moonlight )

oban scientists discover moonlight winter ocean travellers ()

moonlight migration discovered ()

in arctic winter, marine creatures migrate by the light of the moon ()

七星彩走势图带坐标准版moonlight drives marine creatures in arctic winter ()

zooplankton migrate by moon light during arctic winter ()

七星彩走势图带坐标准版study finds arctic creatures navigate by the light of winter moon ()

七星彩走势图带坐标准版arctic animals navigate by the light of winter moon ()

七星彩走势图带坐标准版migrazioni al chiaro di luna per lo zooplancton dell'artide ()

in arctic winter, marine creatures migrate by the light of the moon ()

moonlight drives marine creatures in arctic winter ()

七星彩走势图带坐标准版månesyke for plankton i arktis ()

in arctic winter, marine creatures migrate by the light of the moon ()

in arctic winter, marine creatures migrate by the light of the moon ()

七星彩走势图带坐标准版polar moondance: decades of acoustic data reveal surprising migration patterns of zooplankton in the arctic winter. ()

七星彩走势图带坐标准版el zooplancton del ártico migra bajo la luz de la luna ()

moonlight drives marine creatures in arctic winter ()

in arctic winter, 'werewolves of the deep' hunt by moonlight ()

moonlight drives marine creatures in arctic winter ()

moonlight drives migration of marine animals in arctic winter ()

七星彩走势图带坐标准版sams scientists discover moonlight driving ocean ‘werewolves’ to gather for arctic ocean winter odyssey ()

forskning viser at arktiske dyreplankton er påvirket av fullmånen ()

plankton i arktis kan bli viktig for framtidas klimamodeller ()



七星彩走势图带坐标准版dunbeg primary school. (2016). the science of the arctic ocean.

七星彩走势图带坐标准版university of the third age (u3a), oban, (2016). the rhythms of life.

七星彩走势图带坐标准版interviews with: new york times, new scientist, csmonitor, (2016) “moonlight drives ocean-basin scale mass vertical migration of zooplankton during the arctic winter”.

video abstract for the journal current biology: “moonlight drives ocean-basin scale mass vertical migration of zooplankton during the arctic winter” (2016). .

七星彩走势图带坐标准版university of the third age (u3a), sams (2014). the sams underwater observatory.

association of scottish shellfish growers (assg), oban (2014). calcium in a changing environment.



i review for a variety of journals i.e. chronobiology international, frontiers in bioscience, invertebrate reproduction and development, journal of experimental marine biology, journal of molluscan studies, journal of the marine biological association, marine biology, marine pollution bulletin, polar biology, proceeding of the royal society, scientia marina, soil biology etc.



七星彩走势图带坐标准版member of the society for research on biological rhythms

member of the uk deep-sea sig

member of the challenger society

member of the institute of biology



七星彩走势图带坐标准版arctic ocean cruises aboard the r/v helmer hanssen annually 2014-2017


2015-present senior lecturer in marine biology and chronobiology. sams

七星彩走势图带坐标准版2010-15 lecturer in marine biology and deputy head of ecology department. sams

2006-10 research fellow (arc), sams. research focus: marine chronobiology, sediment impacts on macroinvertebrates, whale-fall fauna ocean acidification.

七星彩走势图带坐标准版2006 research / teaching fellow. university of newcastle upon tyne, uk. responsibilities include: undergraduate teaching, specifically boatwork teaching; collating outputs from nerc egt programme (see below), grant submission and paper generation.

七星彩走势图带坐标准版2002-05 research associate (nerc). university of newcastle upon tyne, uk. part of a collaborative nerc environmental genomics thematic programme investigating the behavioural and molecular basis for tidal rhythms in the marine polychaete nereis virens. work on nereis includes characterization of behavioural phenotypes, cdna library creation, microarray spotting / hybridization and probing for conserved clock genes.

2001-02 Researcher. University of Newcastle upon Tyne, UK.  This period allowed scientific focus on paper writing, grant applications and continuing research into the chronobiology of the polychaete Nereis virens.

七星彩走势图带坐标准版2001 self-employed environmental consultant. consultancy work primarily for e.r.t. caspian which entailed an environmental impact assessment in azerbaijan for british petroleum ltd. this involved benthic mapping and ground-truthing of caspian sea benthic flora and fauna.



2000 PhD 'PhD 'Photoperiodism in the semalparous polychaete Nereis virens七星彩走势图带坐标准版 (Sars)'. University of Newcastle. Supervisor: Peter JW Olive

七星彩走势图带坐标准版1994 msc fisheries biology and management. (supervisor: ernest naylor)

1993 bsc (hons) biology. university of portsmouth

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